Everything about Acoraceae totally explained
Acorus is a
genus of
monocot flowering plants. This genus was once placed within the family
Araceae (aroids), but more recent classifications place it in its own family
Acoraceae and order
Acorales, of which it's the sole genus of the oldest surviving line of monocots. The exact relationship of Acorus to other monocots, however, is still debated by scientists. Some studies indicate that it's placed in a lineage (the order Alismatales), that also includes aroids (Araceae),
Tofieldiaceae, and several families of aquatic monocots (for example, Alismataceae,
Posidoniaceae). Common names include
Calamus and
Sweet Flag. It is known as
vasambu in
Tamil language.
The name 'acorus' is derived from the Greek word 'acoron', a name used by
Dioscorides, which in turn was derived from 'coreon', meaning 'pupil', because it was used in
herbal medicine as a treatment for
inflammation of the
eye.
The genus is native to
North America and northern and eastern
Asia, and
naturalised in southern Asia and
Europe from ancient cultivation. The known wild populations are
diploid except for some
tetraploids in eastern Asia, while the cultivated plants are sterile
triploids, probably of
hybrid origin between the diploid and tetraploid forms.
Taxonomy
Although the family Acoraceae was originally described in 1820, since then
Acorus has traditionally been included in Araceae in most classification systems, as in the
Cronquist system. The family has recently been resurrected as molecular systematic studies have shown that
Acorus isn't closely related to Araceae or any other
monocot family, leading plant systematists to place the genus and family in its own order. This placement currently lacks support from traditional plant morphology studies, and some taxonomists still place it as a subfamily of Araceae, in the order
Alismatales.
The
APG II system recognizes order Acorales, distinct from the Alismatales, and as the
sister group to all other monocots.
Characteristics
These grasslike evergreen plants are
hemicryptophytes, (for example perennial plants of which the overwintering buds are at the soil surface) or
geophytes (for example the overwintering buds are found underground, usually attached to a bulb, corm, tuber, etc.). Their natural habitat is at the waterside or close to marshes, often found with reedbeds.
The inconspicuous
flowers are arranged on a lateral
spadix (a thickened, fleshy axis). Unlike aroids, there's no
spathe (large bract, enclosing the spadix). The spadix is 4-10 cm long and is enclosed by the foliage. The bract can be ten times longer than the spadix. The
leaves are linear with entire margin.
The parallel-veined leaves of some species contain ethereal oils that give a sweet scent when dried. Fine-cut leaves used to be strewn across the floor in the Middle Ages, both for the scent, and for presumed efficacy against
pests.
Species
In older literature and on many websites, there's still much confusion, with the name
Acorus calamus equally but wrongfully applied to
Acorus americanus.
The genus includes as many as six
species:
- Acorus americanus (Raf.) Raf. (formerly known as A. calamus var. americanus) - American Sweet Flag; fertile diploid (2n = 24); occurring in Alaska, Canada and northern USA. Diploid plants in Siberia and temperate Asia may also belong here, but have not been fully investigated ((External Link
)). Recently recognised as a distinct species by the Flora of North America.
- Acorus calamus L. - Common Sweet Flag; sterile triploid (3n = 36); probably of cultivated origin. It is native to Europe, temperate India and the Himalayas and southern Asia, widely cultivated and naturalised elsewhere.
- Acorus gramineus Sol. ex Aiton - Japanese Sweet Flag or Grassy-leaved Sweet Flag; fertile diploid (2n = 18); occurring in the Himalayas to Japan, Myanmar, Thailand, the Philippines.
- Acorus triqueter Turcz. ex Schott (syn. A. calamus var. angustatus) - fertile tetraploid (4n = 48); occurring in eastern Asia, Japan and Taiwan.
- Acorus latifolius Z.Y.Zhu : native to China
- Acorus xiangyeus Z.Y.Zhu : native to China
Acorus from Europe,
China and Japan have been planted in the
United States.
Chemistry
Both triploid and tetraploid calamus contain
asarone, but diploid doesn't contain any. Asarone is a possible precursor in the synthesis of the psychedelic phenethylamine TMA-2. The chemical itself is reportedly without activity at doses up to 70 mg.
Regulations
Calamus and products derived from calamus (such as its oil) were banned in 1968 as food additives and medicines by the
United States Food and Drug Administration.
Usage
Calamus has been an item of trade in many cultures for thousands of years. Calamus has been used medicinally for a wide variety of ailments.
In antiquity in the Orient and Egypt, the rhizome was thought to be a powerful aphrodisiac. In Europe
Acorus calamus was often added to wine, and the root is also one of the possible ingredients of
absinthe. Among the northern Native Americans, it's used both medicinally and as a
stimulant; in addition, the root is thought to have been used as an
entheogen among the northern Native Americans. In high doses, it's
hallucinogenic; Calamus has been used as a "street drug alternative".
Cultural symbolism
The calamus has long been a symbol of male love. The name is associated with a Greek myth:
Kalamos, a son of the river-god
Maeander, who loved
Karpos, the son of
Zephyrus and
Chloris. When Karpos drowned, Kalamos was transformed into a reed, whose rustling in the wind was interpreted as a sigh of lamentation.
The plant was a favorite of
Henry David Thoreau (who called it sweet flag), and also of
Walt Whitman, who added a section called
The Calamus Poems, celebrating the love of men, to the third edition of
Leaves of Grass (1860). In the poems the calamus is used as a symbol of love, lust, and affection. It has been suggested that the symbology derives from the visual resemblane of the reed to the erect human penis.
The name
Sweet Flag refers to its sweet scent (It has been used as a strewing herb) and the wavy edges of the leaves which are supposed to resemble a fluttering flag.
Etymology of the word Calamus
Cognates of the
Latin word
Calamus are found in both
Greek (
kalamos, meaning "reed") and
Sanskrit (
kalama, meaning "reed" and "pen" as well as a sort of rice) — strong evidence that the word is older than all three languages and exists in their parent language,
Proto-Indo European. The
Arabic word
qalam (meaning "pen") is likely to have been borrowed from one of these languages in antiquity, or directly from Indo-European itself.
From the Latin root "calamus", a number of modern English words arise:
calamari, meaning "squid", via the Latin calamarium, "ink horn" or "pen case", as reeds were then used as writing implements;
calumet, another name for the Native American peace pipe, which was often made from a hollow reed;
shawm, a medieval oboe-like instrument (whose sound is produced by a vibrating reed mouthpiece);
chalumeau register, the lower notes of a clarinet's range (another reed instrument)
References and external links
Govaerts, R. & Frodin, D.G. (2002). World Checklist and Bibliography of Araceae and Acoraceae. 1-560. The Board of Trustees of the Royal Botanic Gardens, Kew.
Flora of North America: Acorus
Flora of North America: Acoraceae
Acorales in Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 7, May 2006.
NCBI Taxonomy Browser
Acoraceae in L. Watson and M.J. Dallwitz (1992 onwards) The families of flowering plants: descriptions, illustrations, identification, information retrieval. Version: 27 April 2006. http://delta-intkey.com.
Taxonomy and distribution of Acorus in Maine
(External Link)
Further Information
Get more info on 'Acoraceae'.
|
External Link Exchanges
Do you know how hard it is to get a link from a large encyclopaedia? Well we're different and will prove it. To get a link from us just add the following HTML to your site on a relevant page:
<a href="http://acorus.totallyexplained.com">Acorus Totally Explained</a>
Then simply click through this link from your web page. Our crawlers will verify your link, extract the title of your web page and instantly add a link back to it. If you like you can remove the words Totally Explained and embed the link in article text.
As long as your link remains in place, we'll keep our link to you right here. Please play fair - our crawlers are watching. Your site must be closely related to this one's topic. Any kind of spamming, dubious practises or removing the link will result in your link from us being dropped and, potentially, your whole site being banned. |
